Socially Normative Agency
In Chapter 6 of The Evolution Of Agency by Michael Tomasello we come to human beings. Tomasello says that we humans build on the agency of lizards, squirrels and great apes discussed in earlier posts.
Let’s start with an illustration. Occasionally a group of chimpanzees will hunt and kill monkeys. One chimpanzee will spot the opportunity and scream and start chasing it. Others join in the chase. If they kill the monkey they all try to grab as much as possible. They do not distinguish between those that hunted and free-riders. Only the strongest and wiliest eat.
On the other hand, children as young as three years old act differently. Confronted with a task that requires two actors, they will form an agreement to work together. If they succeed at the task, they split the reward evenly almost always. If one defects from the work or tries to grab the whole reward, the other complains and that is usually enough to bring the offender into compliance. Children who don’t participate in the work are rarely given a share of the reward.
Tomasello says this is a form of agency in humans not present in other species. He calls it socially normative agency.
Evolution of socially normative agency
Tomasello says that hominids, the predecessors of our species, split off from the great apes about six million years ago. They began hunting collaboratively about one million years ago. He suggests that collaboration might have begun with scavenging meat off carcasses. Hominids were smaller and weaker than other species competing for the carcass. They would have had a better chance of success if they worked together, some keeping other scavengers away and some gathering the spoils, then splitting it up among all of the group.
Later, two of our ancestors might agree to work together to hunt large game, agreement would be established, and the two would work together and share the rewards. Over time, collaborative hunting became necessary to meet the demands for food. By about 400,000 years ago collaborative hunting was an established practice. Our ancestors became interdependent, unable to survive on their own.
Psychological processes and feedback control mechanisms
Tomasello says the central mechanism of control in all species is the feedback loop. In the simple form, a lizard uses its perceptionto look for prey, moving on if there is none. In small mammals like squirrels, there is an additional level of psychological control, an executive tier, that enables the individual to keep track of its goals (foraging, burying nuts for the winter, staying ahead of predators etc. They can chanage their behavior if needed for safety or for a better chance of successful foraging. They can choose among alternative actions for reaching those goals.
In great apes, there is third tier, the reflective tier, that monitors the executive tier, and can interrupt the action of the individual to solve problems preventing the individual from reaching its goal, Doing this requires mental processes Tomasello calls rational because they are logically structured.
In the example of three-year olds cooperating in a task, the two reach agreement on joint action. That agreement creates a a new entity, the joint agency. There feedback control mechanism runs each child to the other. Both are entitled to enforce the requirements of the joint agency. If one child stops working, leaves, or refuses to share, for example, the other remonstrates in normative terms. The other will say something like: we don’t do that, or you have to share, or that’s not fair. The offender recognizes that this is a legitimate complaint, and most ofteb changes behavior. Tomasello says this is because each child recognizes and accepts the demands of the joint agency and the right of each to enforce it.
In great apes, the feedback control mechanisms are internal to the individual. In socially normative agency, the feedback loop is in the “we” created by the agreement, and is exercised by both parties as the project proceeds. The feedback control also covers the efforts of the two to collaborate in the task.
Evolutionary changes
The evolution of agency in humans requires greatly increased complexity in the brain: more and better neurons with more and better connections, and a large increase in the pre-frontal cortex, which we think is the location of executive action.
These physical changes were needed for collaboration. Control by agreed normative rules required three new kinds of psychological processes: a) creation of the joint goal: b) allocation of roles; and c) collaboration in self-regulation during the action. Each of these in turn require a chunk of mental processing, This is something humans do automatically, and other creatures don’t.
Those who didn’t cooperate, who didn’t communicate, and who didn’t share were left behind, without mates, and without offspring. The mental processes that enable collaboration, including collaboration, became targets of natural selection, probably in a form of co-evolution.
Cultural groups differentiated by their norms
Collaborative hunting and reward sharing was very successful for our ancestors. By about 150,000 years ago, human population had grown to the point that we began to live in larger and larger groups. Successful hunting of larger game was just one part of survival. Others in the group foraged for plants and small game, some became adept at creating tools snd weapons, others at weaving and making clothes, others learned about curative herbs and healing and so on.
Tomassello says that our ancestors were living in bands loosely connected to each other by cultural norms related to diet, stone knapping, languages, foraging techniques and more. Membership in a cultural group became necessary for survival as these cultural groups competed for resources. Again, those who couldn’t manage the psychological processes necessary for participation in the cultural group were shunned and didn’t reproduce. Compliance with cultural norms became a target of natural selection.
Cultural groups survived by teaching their offspring proper, that is normative, behaviors and techniques. Long childhoods and adolescence were periods of training and learning the social norms of the cultural group. In operation, these cultural groups made cpllective decisions, and enforced them. Tomasello calls this cultural agency. The feedback control is operated by the entire group.
Here’s how Tomasello describes the evolutionary process up to about 100,000 years ago:
… modern human brains are three times larger than those of other great apes, with an expanded prefrontal cortex (the main seat of executive functioning) and insula (the main seat of social emotions), and are structured by unique types of neurons with more complex dendritic structures Consistent with this analysis, González-Forero and Gardner … find that much of the brain growth characteristic of modern humans during this period was concerned with adaptations for cooperative interactions and cultural learning. P. 106-7; cites omitted.
Discussion
Regardless of your view of Tomasello’s model, one thing is clear from the evidence he cites. Human evolution runs side-by-side with increasing levels of cooperation. Our ancestors taught each other; we literally dragged ourselves up by our bootstraps. Social learning and cooperation are an integral part of our evolution.
In The Dawn Of Everything David Graeber and David Wengrow discuss the Thomas Hobbes theory that early humans fought each other for resources, the war of all against all. That didn’t happen. We should stop talking like it did.
In the same way, the bizarre neoliberal idea that we humans are atomistic utility-maximizers struggling against each other to get all we can is madness. Any academic who takes this view should be laughed out of their profession.
I’ll have more to say about this in the next post.
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Thank you.
Great post! Socially normative agency is a great term for something I have observed without having the vocabulary necessary to describe.
One small personal observation. I believe I have observed this behavior in children even younger than three. My youngest two children are identical twins. As soon as they could walk, I saw them develop this sort of behavior. Specifically, they discovered that if one of them would take off toddling as fast as he could, Dad would give chase. The other boy would use that opportunity to open a drawer or closet that was off-limits, take out two things and hand one to his brother when he came back around (our house at the time had a loop – living room-entry way-dining room-kitchen-living room).
I may have encouraged that behavior by not chasing as fast as I could. They invented it all by themselves.
Just out of curiosity. Was it always the same child who was the fleeing decoy, or did they sometimes switch roles?
As I recall, they would switch off.
I have 2 brothers that are twins, the oldest of 4 brothers. As a career biologist, it has been both fascinating and depressing to watch these 65+ year old men revert back to their childhood co-dependencies…
That is a delightful illustration! Thanks.
I have seen similar with the interactions of my three granddaughters. A 9 year and 7 year old who are sisters and the third a cousin of 3 years old.
Of course as grandparents we do not have them all the time – fortunately we can spoil them and then give them back to their respective parents.
Be careful with the bootstrap language. I think what you are saying here is that we pulled one another up by our bootstraps, which runs completely counter to the more usual phrase “pull yourself up by your *own* bootstraps” which is language that evokes the mythical Rugged Individual.
Thank you for the bootstrap counsel/addendum.
Quote: “Tomasello says this is a form of agency in humans not present in other species. He calls it socially normative agency.:”
Tomasello makes that assertion and all that follows relies on everyone else agreeing his assertion is both factual and final, that no other species acts as some humans do, …but I’m not so sure of the primacy of his assertion..
Tomasello’s example is simply a form of adaptive cooperative behavior. Two humans showing cooperative behavior limited to or confined by certain factors.
Just a single example of cooperative adaptive behavior patterns we are starting to recognize as the more prevalent behavior in life forms on all levels, even right down to the cellular level, this as opposed to the old belief that evolution is primarily the result of tooth and claw or other forms of competition.
Cooperation is the more central driver of evolution.
Example: Vampire bats hunting failure rate is very high, missing two days of a blood meal is fatal to a vampire bat, but vampire bats have evolved to share blood meals. And those who don’t share get a reputation and are excluded from the sharing. Without sharing, four out of five bats in a given colony would die. Adaptive cooperative behavior.
Complex adaptive cooperation is able to be observed more readily now that we have computers processing ever expanding quantities of video observations to help us define adaptive behaviors to a degree previously unattainable by a limited number of sole source single observers.
Computer simulations are revealing a trove of behavioral cooperation we never gave a thought to in the not too distant past. From human behavior, to animals, insects right down to the cellular and molecular level.
We are also learning that complex adaptive behavioral cooperation between cells is a widespread phenomenon in nature, from microbial populations to multicellular organisms.
What we know of the cooperative behavior in life’s other forms is not able to be defined with such finality as Tomasello would have you believe.
There’s no argument that humans have learned to expand their adaptive cooperative behavior, …but so have other species, as well as have other lifeforms.
Contriving an ‘agency’ that humans alone can or will attain is simply another attempt to elevate humans above other life forms. To place humans outside the systems, networks and ecology that all known life shares.
This is a very helpful comment, thanks. I agree that adaptive cooperation is everywhere. A good example is in the documentary Fantastic Fungi, showing the cooperation among fungi; it’s a great movie as well as an informative one.
I read Tomassello’s book as laying out a model to help us see how agency evolved. The key point is that in other species cooperative behaviors are controlled by internal structures of the individual brain. Tomasello claims that human brains evolved in ways that enabled greater and greater cooperation by setting up feedback control loops among members of our species. This is from the beginning of Chapter 6:
Tomasello says the brains of other species did not evolve to support a kind of general cooperation as our species did. Vampire bats may share a blood meal, but they don’t hunt together, which perhaps would be a more useful way of insuring survival. And they certainly did not evolve to discuss and agree to work cooperatively, or to structure a society that depended on cooperation.
Maybe we should think of this as a genealogy of cooperation as much as a model of the evolution of agency.
Quote: “Tomasello says the brains of other species did not evolve to support a kind of general cooperation as our species did.”
That’s a handy statement to maintain his belief that we humans are ‘superior’ to other life forms.
And your quote is a similar unproven assertion: “…they certainly did not evolve to discuss and agree to work cooperatively, or to structure a society that depended on cooperation.”
I’ll point to the cooperative behavior of ants or bees and point out that until we can communicate with them, (or bats), we can’t know what’s in their minds or what they communicate amongst themselves.
As to human’s ‘success’ through our supposed ‘superiority’, biodiversity holds the key to the stability of our habitat and ultimately life itself, we call ourselves the most intellectual beings to have evolved but through our willful actions we have done more damage to biodiversity than any other life form, and despite our large cranial capacity, we’re still heading headlong towards our own extinction.
We can tell ourselves we are the superior life form, but the proof is gonna be a different matter.
There’s nothing about superiority in my discussion. Our species has survived. That’s success.
One way to think about it is that bats are unquestionably superior to rats when it comes to flying, just as rats are superior to bats when it comes to digging, but neither statement should be taken as a judgment the mobility of one species on the whole versus another.
There is absolutely no question that humans have a number of mental capabilities that exceed any other species, but likewise we shouldn’t assume that means in all facets we exceed every other species.
Thanks Ed, for this discussion and others you’ve hosted.
It wasn’t long ago the generally accepted belief was that fish couldn’t feel pain, that biologically their brains and nervous systems were not considered ‘developed’ enough to process such basic stimuli as pain. Today we know better.
There is evidence today that even plants can respond to stimulus, (pain), and communicate that to other plants. We have also learned that trees can help feed other trees that demonstrate signs of stress. Anyone expressing that belief not long ago were summarily roundly ridiculed.
Feedback loops certainly aren’t uniquely human.
When Tomasello speaks of ‘human domination’ of other species, I can’t help but see his belief as comparable to him expressing ‘superiority’.
As long as our awareness of reality is evolving, the Tomasellos among us have to accept that not everyone is going to go along with assertions not backed up with proofs.
It may comfort some to think humans have powers beyond the reach of other life forms, ‘authorities’ appearing in many forms have long preached of human ‘dominion’ over nature.
Like as not, that belief may be false, and it may very well be such belief could ultimately lead to our self manufactured downfall.
Great points, AR, and thanks for the continued posts on this topic, Ed.
Jeremy Bentham: “The question is not, Can they reason?, nor Can they talk? but, Can they suffer? Why should the law refuse its protection to any sensitive being?”
This recent US Supreme Court case was about animal welfare rather than animal rights, but was the first to recognize that animal welfare is more important than the pursuit of interstate commerce: http://www.humanesociety.org/news/animal-welfare-supreme-court
And the more we learn, the closer the US comes to recognizing the rights of non-human animals, which have been recognized at the State Supreme Court level—most recently Chimpanzees Tommy, Kiko, and Cecilia (www.nonhumanrights.org/blog/cecilia-chimpanzee-legal-person/)—but remain unrecognized by the US Supreme Court.
As an animal activist, my most fervent hope is to see them recognized before I shuffle off this mortal coil.
*EDIT* Mods: I tried 3 times to remove the http for the first link, but it just kept coming up. Sorry.
Thank you for this, the concept has been like a splinter in my mind since I read it in the OP. It amazes me that we have always shared spaces with animals, domesticated or otherwise, but the conventional wisdom is that those organisms are half-wits.
Dr. Tomasello is certainly an authority, but “not present in other species?” Recently I read “The Mind of a Bee” by Lars Chittka, and it goes into quite some detail about how bees think, make decisions, and cooperate for the good of the hive (in the case of hive bees) or nest (bumblebees), as shown by rigorous studies and experiments at least since Karl von Frisch’s work around 1927. Even in these supposedly “lower” animals; preservation of a species –- whether “complex” or “simple”— may require learning and consciousness as well as hard wiring, and require cooperation. The question for me is whether humans are actually cooperating to preserve our species any better than insects. Current events make me wonder.
As I understand it, Tomasello isn’t just talking about cooperation (which occurs in many species), but cooperation to obtain a reward that is then shared fairly equally among those who contributed, with norms enforced if an individual attempts to take more than their fair share / attempts to obtain a share despite not having contributed to the effort. If you’re suggesting that bees do that (your query “not present in other species?”), would you give an example of what kind of reward bees cooperate to obtain, and how they respond if a bee that didn’t participate attempts to share in the reward?
Well, we can’t push an analogy too far, but (perhaps unlike humans) insects don’t act out of arbitrary selfishness; they occupy specific “professions.” In hive bees, a queen never forages; she is fed, but mates and provides the eggs to perpetuate the colony. The famous “bee dance” is an example of bees communicating to tell the other foragers where the good nectar sources are or, when swarming, where a good possible new nest location may be found. They don’t conceal their sources.
I wouldn’t call it an analogy, but a set of criteria re: what constitutes “socially normative agency.” And I’m asking if there’s evidence of bees demonstrating that, or if they’re instead demonstrating a different kind of cooperation that’s found in a number of species.
I don’t know, haven’t read Tomasello’s book. But I do question 1) whether humans are alone in this agency and 2) that it works to the benefit of human survival, as others here note. It seems to me likely that insects will survive far longer than homo sapiens.
“…what kind of reward bees cooperate to obtain, and how they respond…”
Bees are actually a textbook case of reward systems: foraging sites, individual and colony hygiene, for example. The worker progression from baby-tender to forager as they age. Whether there is a conscious decision making process or hard coded in the DNA might be a distinction without a difference. After all we’re just beginning to scratch the surface here.
I, too, read that book. Lots of fascinating stuff in there.
I had same reaction as you reading the “not present” line. My mind first went to the pair of robins tag team feeding their youngsters in the nest on my back porch this past spring. Conveniently viewable from kitchen window.
Then to an account by a woman who a crow befriended by continually dropping off all manner of gifts for her — anything shiny and/or brightly colored. Even someone’s lost ring one day.
Then to Lars.
Besides “that looks interesting” aspect, I read the bee book partly to avoid reading political/historical horror stories for awhile.
Not a complete escape. Lars writes of key bee guys teasing out remarkable things decades back in very trying times: A black guy in U.S.; a Jewish guy in Europe. Sigh…
Thanks for this helpful illustration. AlaskaReader above makes a similar point; please see my reply. In a comment on an earlier post I quoted this from Tomasello:
I’m not sure I’d agree that insects think, even collectively. Maybe I’m prejudiced by something Richard Feynman wrote in Surely You’re Joking Mr. Feynman, quoted here: https://theswarmlab.com/blog/2015/08/08/Richard-Feynman-had-a-thing-for-ants/
You might enjoy Chittka’s book. It has fascinating illustrations. He describes many experiments that show how bees think and how they adapt to circumstances. Admirably and unusually, he gives plenty of credit to key researchers in that field, as FeistyBlueBird points out. Plus yes! A wonderful world-disasters-avoidance read.
“Those who didn’t cooperate, who didn’t communicate, and who didn’t share were left behind, without mates, and without offspring.”
Might this also be true of the 21C sub-grouping, known as ‘incels’? /s
Thanks for your analysis. It encouraged me to purchase this book.
Anyone who has worked on a large project knows that cooperation among the group working on the effort is an essential element to the project’s success. As a technical person, I have worked and managed a number of projects over my lifetime and one of the major challenges is to form a team of people who not only have the correct skills, but who can also cooperate with others.
Unfortunately, and for reasons I do have never understood, modern society tends to award credit for successful outcomes only to a few people, often only one person.
For example, Elon Musk is viewed by many in our society as the genius who single-handedly created Tesla and Space-X.
This is nonsense. These companies were already in existence and doing well before Elon stepped in. What he can perhaps be credited for is that he pumped in capital for them to grow, as well as mostly leaving them alone to prosper except for his occasional destructive actions. In fact, in my view these companies are successful despite Elon being part of them and that is probably due to the capabilities and cooperation among the others at these organizations.
As another example, Thomas Edison has always been viewed as one of the greatest inventors of all time. I don’t doubt that he had many great ideas, but the number of patents credited to him is ridiculous to anyone who has ever made any patentable inventions. There is just not enough time in a person’s life to make a sufficiently worthy contribution to be declared as an inventor on that number of patents.
We also see similar kinds of credits given to sport stars wherein one member of the team is viewed as the reason for any success of the team.
No matter how much one or a few people may have contributed, it is almost always the case that the others collectively contributed far more to these organizations that one person or a few people ever could.
Not only do I wish I understood why this happens, it leads to a number of questions such as:
Is this a new thing among humans or has it always been present?
What is the impact of this on our ability to cooperate? To me, it would seem to be negative.
Is this a good or bad thing for our society?
There’s an excellent description of the overhead that Elon Musk imposes on any organization he is involved with.
The first paragraph is:
“Back when I was at SpaceX, Elon was basically a child king. He was an important figurehead who provided the company with the money, power, and PR, but he didn’t have the knowledge or (frankly) maturity to handle day-to-day decision making and everyone knew that. He was surrounded by people whose job was, essentially, to manipulate him into making good decisions.”
You can read it at https://www. reddit.com /r/EnoughMuskSpam/comments/z2ofwk/i_was_an_intern_at_spacex_years_ago_back_it_when/
The URL has a space and is folded onto 2 lines to make it unclickable, following site policy.
Maybe a different way to think of the “one person most important” question is to think of the given person or persons as a catalytic agent without which the group effort directed toward a certain goal would not have happened. So, yes, for a given large corporate project the team that completes it is obviously dominant, but the project wasn’t happening without some sort of agency bringing it into existence. Edison, if the patents weren’t all his, most likely was the catalytic agent for those that did invent whatever and patent under his name. Those catalytic agency types seem to be the ones that get the most credit.
In relation to that catalytic process, I suspect the concept that man records, thinks, and communicates ideas through different languages and times, and the man can use those ideas as a tool to propel her/himself forward (I have seen so far because I have stood on the shoulders of giants) to more advanced ideas is more a source of evolution than cooperation. Evolution is about change. Cooperation is about, well, cooperation.
In relation to reasons for physical growth of the brain Frank Wilson had a book some years ago titled The Hand. I haven’t re-read it in some time but I seem to remember that he correlated the growth of the brain with physical changes in the hand, necessary to gain the the ability to throw a spear among other things. Again it’s misty in my memory but I seem to remember him using the concept of necessary growth in the brain to accommodate uses of the hand. As examples we all have played catch and have placed our hand in a position to catch an object without looking at it, juggling, tying a knot behind one’s back, playing a piano. It’s quite a fascinating book. It may work as a theory in line with Tomasello’s reasons for brain growth.
this is my theory about the extinction of the Neanderthals. All evidence seems to indicate that they did not form large social groups, and my theory is that they did not have the psychological mechanisms to engage with larger groups than close family members.
Interesting.
A good chunk of the scientific community doesn’t necessarily believe the Neanderthals went extinct, but that they were subsumed or absorbed by Homo sapiens via interbreeding, as evidenced by Neanderthal genes in our present population.
This theory/study is interesting:
“Dreaming may have evolved as a strategy for co-operative survival” -David Samson, 12/20/23
https://theconversation.com/dreaming-may-have-evolved-as-a-strategy-for-co-operative-survival-216328
Another book on this topic worth reading is Edward Wilson’s The Social Conquest of Earth. He also discusses the evolution of humankind along the lines of the need for/advantage in cooperation. Sadly we’re seeing a devolution nowadays into a form of tribalism that is self-defeating in the long run and doesn’t bode well for the prospects of the human race.
“…we’re seeing a devolution nowadays…”
“Nowadays?” When has it ever *not* been that humans’ propensity for large scale cooperation has produced as devastating results via acts of predation against our own species as compared to species-enhancing results from cooperative behaviors.
Yes, we’re almost universally connected with each other these days (and that took unthinkable amounts of cooperation), but those connections are as volatile and pernicious, by and large, as they are stability-enhancing and beneficial. And the largest species-threatening issue we’re faced with right now (other than the ubiquitous, civilization-ending nuclear war one) is the environmental one that has intra-species competition between the haves and have-nots driving us pall-mall to a mass-extinction and societal-collapse hellscape.
It’s comforting to think that the lessons we learned hunting big game on the savannas and tundras — or cooperatively and permanently altering our local environments in the service of human domination of them — will carry us around or beyond those increasingly likely catastrophic results, but that’s certainly not a rational or honest appraisal.
Are we well and truly fucked? I have no actual idea, but rose colored glasses will not provide any answers or solutions to that question.
I think my point is that with the technological advances we as a species have made also comes the danger of destroying ourselves in any number of ways (both the quick and the slow). I didn’t think I was wearing rose-colored glasses at all. In fact I’d count myself among the pessimists. Until humans recognize that we are all in this together, we’re in trouble. And the voices screaming loudest nowadays command a global audience and I don’t hear them preaching peaceful cooperation. The old saying goes “A rising tide lifts all boats”…but what about those who don’t have boat? Either we welcome them aboard and maybe build some more boats or we’re going to end up fighting over who doesn’t drown.
It’s been a very long time since I read Richard Dawkins’ The Selfish Gene*, but I think it may have something to say about other species besides humans acting in altruistic fashion in order to ensure the success of a specie’s genes.
Tomasello places a lot of emphasis on humans as differentiated from other species, but it’s rather like American exceptionalism or white supremacy — what the hell is he basing this emphasis on when many other species act collaboratively and collectively to assure the ultimate survival of their kind?
There’s an issue of qualia here we can’t transcend. Qualia is one’s unique experience which can’t be conveyed with absolute certainty, ex. when I say something is red, you have an idea of what I mean by the color red but there’s no way to be certain both of us have the same experience of that color. This is the problem with assumptions about consciousness and intelligence in other species: as humans we make a lot of guesses about other species but we’re doing so based on the use of human-selected tools measured against human-identified references. What if there is far more collaboration and collective action in other species but we puny humans cannot grasp it with our frames, unable to share other species’ experience of reality?
This recently published study about bonobos also tweaks Tomasello’s use of chimapanzees:
https://www.sciencenews.org/article/bonobos-humans-cooperation-evolution
I dunno’. I suspect even plants have some form of cooperation and collaboration based on what we’ve learned about their ability to communicate; what constitutes agency in their frame?
Inter-plant communication through mycorrhizal networks mediates complex adaptive behaviour in plant communities (2015)
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4497361/
Aboveground mechanical stimuli affect belowground plant-plant communication (2018)
https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0195646
* yes, I know, that Dawkins, the problematic one. Still need to consider what he’s written about altruism as a benefit to genetic success.
Thanks! And the work of Robin Wall Kimmerer.
I see altruism, and other traits, working this way. Individual members of a species express varying levels of altruism. To the extent that altruism increases an individual’s survivability, and continues to do so as the immediate environment changes over time, then individuals with higher levels of that trait would become a growing percentage of the members comprising that species.
Selective pressure – speedier dying off of individuals with traits outside a useful norm – would keep altruism within a given range, as expressing too much or too little would shorten the lifespan of an individual, without a corresponding benefit, individually or at the species level.
At the species level, if altruism increases the survivability of other members of the species, the percentage of members acting altrustically would increase, but more slowly than if altruism directly increased an individual’s personal survival.
Socially, we can see how traits such as altruism are open to abuse by predators internal to the species, which helps them survive and proliferate.